A decade has passed since the events of Angelology when Verlaine saw Evangeline alight from the Brooklyn Bridge, shocking him with the sight of her new wings. Verlaine is now an elite angel hunter for the Society of Angelology, dedicated to killing all of Evangeline's kind. He realises she is nothing like the other angels and when she is abducted in front of him by the society's most wanted criminal he sets of on a chase. He follows the pair from Paris to Russia and deep into Siberia where the truth of Evangeline's origins awaits, along with forces that could destroy them.