The Klamath River flows about 255 mi (410 km) from the outlet of Upper Klamath Lake through southern Oregon and northern California to the Pacific Ocean. The first 21 mi of the river, just downstream of Upper Klamath Lake, are bounded by Link River Dam and Keno Dam (fig. 1). Water quality in this reach has been classified as "very poor" by the State of Oregon (Mrazik, 2007) and was designated as "water quality limited" on Oregon's 303(d) list for exceeding ammonia toxicity and dissolved oxygen criteria year-round, and pH and chlorophyll a criteria in summer (Oregon Department of Environmental Quality, 2007).