The tree recounts its visits over the years by a special human who was born in the same year. As a boy, this human climbs the cottonwood's branches to watch the river and dream. As a father, he brings his daughter to visit. As an old man he grieves to see the tree knocked down but rejoices when he sees new sprouts emerging from the stump. He knows the tree is not done yet!

Captions throughout the book relate the natural histories of animals in and around the tree, and the backmatter offers further resources.