As excitement and the scent of cupcakes filled the air at Genevieve Parfait's Patisserie, the tiny Parisian rabbit had no idea that baking for the King's upcoming birthday gala would set off a chain of events that would take her from the Towers of London to the Grand Canal of Venice. When her competitor Jacques, a rat with both a jealous bone and devious mind, steals her sugary masterpiece and plots to keep her as far away from Paris as possible, Genevieve has no time to waste in finding her confection, a tower of irreplaceable cupcakes made of the rarest and most extravagant ingredients. Excitement turns to high anxiety as Genevieve joins forces with the bold and brash Highlander, Collie Duncan MacTartan, a former Scotland Yard detective and his former partner in police work, the proper and particular British Terrier, Finneas Crumpett. As the unlikely threesome follow clues to some of Europe's most famous landmarks, rookie reporter Sophie Felinee follows their trail bringing international attention to the case and the support of all of Europe. In a race against time, Genevieve and her friends discover much more than cupcakes before the mystery ends in a spectacular and raucous conclusion from high atop the Eiffel Tower.