The heroine of The Wings of a Dove is Milly, an extremely rich American girl, who has been left with no living relatives. She spends much of her time with friend and fellow traveler Susan Shepherd Stringham and together they become part of the social circle of Maud Lowder, a friend of Susan's who lives in London. Milly and Susan are not introduced to the readers until the third chapter. The first two chapters are taken up with Kate Croy, who is Mrs. Lowder's niece, and Merton Densher, a journalist whom Kate is in love with.