Makoto Amano wants to become an actor, but his stern father has decreed that the only way Makoto can pursue his dream is to spend the last two years of high school as a girl! He quickly makes friends with popular tomboy Ito Miura, another drama enthusiast at this new high school and the only student to find out his secret - but are they more than pals?

Makoto's doting sister Tsubaki finds out that Makoto hasn't kept his secret from Ito! She threatens to tell Makoto's father unless Makoto and Ito's homeroom can win a school competition. Then Makoto accompanies the entire Miura household to their ancestral home to meet Ito's eccentric, sharp-witted grandmother. Little does Ito know that her past will catch up with her there…literally!