When Gerry Tremble and his family move from Connecticut to Florida to run his ailing father's hotel, they leave behind one family member who isn't about to take it lying down. He was just a pet cat they called Tony, but Windrusher is determined to find his family. Setting off on a perilous journey, Windrusher calls on his connections to a mystical world of cat gods and goddesses to help him reunite with his loved ones.